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Biernaskie and Tyerman (2005) suggest that Whitham et al. (2003) misuse the concept of the extended phenotype (Dawkins, 1982), in which a behavioural trait such as nest construction by birds or dam building by beavers is an adaptive genetic trait of the individual. Dawkins recognizes that for extended phenotypes such as dam building to have evolutionary consequences for beavers, the dam building trait must have fitness consequences for the individuals expressing the trait. However, because there are ecological and evolutionary consequences of extended phenotypes such as beaver dams on other species, it is also important to consider those impacts that do not necessarily involve feedbacks. Even Dawkins recognizes these other effects.