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Apoplastic effectors, in particular, often contain several disulfide bonds [17] and predicted secretomes of pathogenic fungi contain proteins with elevated levels of cysteines compared to all proteins (Fig 1A). [...]the criteria of small and cysteine-rich can be used to mine predicted secretomes for apoplastic effectors and reduce the number of candidates [18,19]. [...]an increase in the number of identified fungal effectors might enable machine learning approaches for unbiased prediction, which could lead to the discovery of protein properties common to fungal effectors.